The paper can be downloaded for free
here.
A 425-Million-Year-Old Silurian Pentastomid Parasitic On Ostracods by David J. Siveter, Derek E.G. Briggs, Derek J. Siveter and Mark D. Sutton,
Current Biology, DOI: 10.1016/j.cub.2015.04.035
Siveter et al, 2015 wrote:Highlights
•A 425-million-year-old pentastomid is the first adult example from the fossil record
•It is the only known fossil pentastomid preserved with its host
•Uniquely among pentastomids, some specimens are attached to the host externally
•Living pentastomids may have originated as ectoparasites on marine invertebrates
Summary
Pentastomids (tongue worms) are worm-like arthropods known today from ∼140 species [ 1 ]. All but four are parasitic on vertebrates. Their life cycle typically involves larval development in an intermediate host followed by maturation in the respiratory tract of a definitive terrestrial host. Fossil pentastomids are exceedingly rare and are known only from isolated juveniles [ 2–6 ]. The identity of the possible hosts of fossil pentastomids and the origin of their lifestyle have generated much debate. A new, exceptionally preserved species, described based on adults from 425-million-year-old marine rocks, is the only known fossil pentastomid associated with a host, in this case a species of ostracod crustacean. The pentastomids are preserved near eggs within the ostracod and also, uniquely for any fossil or living pentastomid, are attached externally to the host. This discovery affirms the origin of pentastomids as ectoparasitic on marine invertebrates. The terrestrialization of pentastomids may have occurred in parallel with the vertebrate invasion of land.